Last update: 2007-01-12 15:34:09

Richard L. Boyce

Northern Kentucky University
Dept. Biological Sciences
Nunn Drive
Highland Heights, KY 41099
Telephone: 859 572-1407
FAX: 859 572-5639


Yale University, Graduate School, Department of Forestry and Environmental Studies, M.S., December 1986; Ph.D., May 1990
Yale University, School of Forestry and Environmental Studies, M.Forest Sci., May 1985
University of Illinois, Department of Astronomy, August-December 1981
Williams College, Astrophysics, B.A. (cum laude, with honors), June 1981

Professional Positions:
  • Associate Professor(August 2006-present), Department of Biological Sciences, Northern Kentucky University, Highland Heights, Kentucky.
  • Assistant Professor (August 2002-July 2006), Department of Biological Sciences, Northern Kentucky University, Highland Heights, Kentucky.
  • Research Affiliate (May 2002-present), Institute of Arctic and Alpine Research, University of Colorado at Boulder, Boulder, Colorado.
  • Assistant Professor (September 1995-August 200), Department of Biological Sciences, University of Denver, Denver, Colorado.
  • Visiting Assistant Professor (September 1994-August 1995), Department of Biology, Swarthmore College, Swarthmore, Pennsylvania.
  • Postdoctoral Research Associate (January 1990-June 1994), Environmental Studies Program, Dartmouth College, Hanover, New Hampshire.

  • Research Interests and Activities:
    Factors driving "bark-stripping" in Rocky Mountain bristlecone pine; canopy uptake and allocation of nitrogen by conifers; ecophysiology of red cedar; the effect of the invasive Amur honeysuckle on forest understory herbs; effects of calcium deficiency on ecophysiology of trees at HBEF, including chlorophyll fluorescence; development of Fuzzy Set Ordination (FSO) for use with plant community data; undergraduate student learning; spatial statistics.

    Distinguished Awards and Memberships:
    Phi Beta Kappa National Honor Society Member, Sigma Xi Scientific Honor Society 1998 Top Professor, University of Denver Mortar Board Society Member, Ecological Society of America Chair, Rocky Mountain Chapter, ESA, 1997-2002 Webmaster, Rocky Mountain Chapter, ESA, 2002-Present Member, Governing Council, ESA, 1997-2002 Member, Program Committee, ESA, 1998-2002 Member, American Association for the Advancement of Science Member, Kentucky Academy of Sciences Secretary, Botany & Microbiology Section, 2004-2005 Chair, Botany Section, 2005-2006

    Selected Publications:

    Boyce, R.L. 2007. Chlorophyll fluorescence response of red spruce and balsam fir to a watershed calcium fertilization experiment in New Hampshire, USA. Canadian Journal of Forest Research in press.

    Boyce, R.L. 2007. Invasive shrubs and forest tree regeneration. Journal of Sustainable Forestry in press.

    Boyce, R.L., J. R. Larson, and R.L. Sanford, Jr. 2006. Phosphorus and nitrogen limitations to photosynthesis in Rocky Mountain bristlecone pine (Pinus aristata) in Colorado. Tree Physiology 26: 1477-1486.

    Boyce, R.L. April 2005, posting date. Life Under Your Feet: Measuring Soil Invertebrate Diversity. Teaching Issues and Experiments in Ecology, Vol. 3: Experiment #1 [online].

    Boyce, R.L., R. Clark, and C. Dawson. 2005. Factors determining alpine species distribution on Goliath Peak, Front Range, Colorado, U.S.A. Arctic, Antarctic, and Alpine Research 37:82-90.

    Tomaszewski, T., R.L. Boyce, and H. Sievering. 2003. Canopy uptake of atmospheric nitrogen and new growth nitrogen requirement at a Colorado subalpine forest. Canadian Journal of Forest Research 33:2221-2227.

    Boyce, R.L., A.J. Friedland, C.B. Vostral, and T.D. Perkins. 2003. Effects of a major ice storm on the foliage of four New England conifers. Écoscience 10:342-350.

    Friedland, A.J., R.L. Boyce, and C.B. Vostral. 2002. Winter- and early spring microclimate of a mid-elevation conifer forest canopy in central Vermont. Agricultural and Forest Meteorology 115:195-200.

    Boyce, R.L., C.B. Vostral, and A.J. Friedland. 2002. Winter water relations of New England conifers at their upper elevational limits. II. Modeling. Tree Physiology 22:801-806.

    Vostral, C.B., R.L. Boyce, and A.J. Friedland. 2002. Winter water relations of New England conifers at their upper elevational limits. I. Measurements. Tree Physiology 22:793-800.

    Boyce, R.L, and P.C. Ellison. 2001. Choosing the best similarity index when performing fuzzy set ordination on binary data. Journal of Vegetation Science 12:711-720.

    Schauer, A.J., A.W. Schoettle, and R.L. Boyce. 2001. Partial cambial mortality in high-elevation Pinus aristata (Pinaceae). American Journal of Botany 88:646-652.

    Co-Authors and Collaborators:
    Clark, Rebecca; Durtsche, Richard D. ; Friedland, Andrew J.; Larson, Jennifer; Long, Andrew; Mackin, Gail; Martines, Kristi; Metzger, Jessica; Perkins, Timothy D.; Russell, K.C.; Sanford, Robert L., Jr. ;Schauer, Andrew J.; Schultheis, Patrick ;Sievering, Herman; Stienitz-Kannan, Miriam; Tomaszewski, Timothy ;Vostral, Chandra B. Walters, Keith

    Graduate Students:
    Joly, Jennifer; Schauer, Andrew J.; Tomaszewski, Timothy; Vostral, Chandra B.

    Graduate Advisor:
    Berlyn, Graeme P.

    Role at the Hubbard Brook:
    Use chlorophyll fluorescence to study Ca deficiency responses

    Title at Hubbard Brook:
    Principal Investigator